A.D.'s thinking about this whole topic was hugely shaped by many conversations with Robert M. May; we are very grateful to him for the insights provided by these discussions and to John Avise, Doug Erwin, Michael Donoghue, Nadia Talhouk, and Alejandra Jaramillo for comments on an earlier draft. The first draft of the chapter was written in Kilimanjaro, Nairobi, and Heathrow Airports; A.D. thanks British Airways and Precision Air for the patience, care, and attention of their ground staff. This work was supported by National Institutes of Health and National Science Foundation Ecology of Infectious Disease Program Grant DEB-0224565.

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