Deciduousness: most leaves are shed in November, but some always remain. New foliage is produced by April, but the trees in closed forests are scarcely deciduous and new leaves appear with the flowers (Savill & Fox 1967)
Dispersal: probably by animals Timing: flowering period: mainly in November to December but can be prolonged to June (Savill & Fox 1967); fruiting period: April to May (Savill & Fox 1967)
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