Habitat

Species occurrence increases with rainfall to reach a very wide optimum range between 1800 to 3300 mm/yr (logistic regression analysis, Chi2 test). It frequently occurs near the coast (Chi2 test). It is almost exclusively found in disturbed habitats such as forest edges, forest gaps, secondary forests, plantations, and along roadsides. Sometimes found along riverbanks and swamps. On sandy or lateritic soils.

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