Grasslands grow in regions with limited annual rainfall (Ehleringer et al. 1997) and possess several modes of function. In the Mediterranean-climate regions of the world, grasslands consist mainly of annual grasses that are functional during the winter and spring growing season and use the C3 photosynthetic pathway. Grasslands in temperate continental regions are perennial and physiologically active during the summer growing season. Often, C3 and C4 grass species coexist, but the proportion of the mixture shifts toward C4 species as one moves south. In the subtropics, most grasses pho-tosynthesize with the C4 pathway.

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