Figure 5 Changes in P/R ratios during ecological (autotrophic) and urban (heterotrophic) succession. Although P/R decreases as the amount of biological material increases during ecological succession, the result is a mature (climax), sustainable community. In contrast, the accumulation of physical material during urban succession frequently results in a fragile, nonsustainable community.
and urban components if societies are to design and implement the concept of sustainability at the agrolandscape scale (see Figure 1). This approach should simultaneously conserve and enhance biotic diversity at greater temporal and spatial scales.
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